There is some truth in the coaching advantages they had during the Spurrier/Meyer eras and even them being in Tennessee's head a little bit in more recent times, but it really all comes back down to Florida's natural recruiting advantage.
I know we recruit some studs and they recruit some studs, but it's those guys in between the studs, the meat of the rosters, where UF has had the distinct advantage, especially when it comes to speed, for as long as I can remember.